Researchers mapped 16,600 theropod footprints at Carreras Pampas in Bolivia’s Torotoro National Park, covering roughly 80,570 sq ft (7,485 m²). The prints, preserved in soft mud from about 101–66 million years ago, include walking, running and over 1,300 shallow-water swimming traces, plus several hundred bird tracks. Variations in print size and spacing reveal a wide range of animal sizes and behaviors and provide behavioral details that bones alone cannot.
World’s Largest Dinosaur Tracksite: 16,600 Theropod Footprints Mapped in Bolivia

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